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Parasitism is a major aspect of evolutionary ecology; for example, almost all free-living animals are host to at least one species of parasite. Vertebrates, the best-studied group, are hosts to between 75,000 and 300,000 species of helminths and an uncounted number of parasitic microorganisms. On average, a mammal species hosts four species of nematode, two of trematodes, and two of cestodes.[78] Humans have 342 species of helminth parasites, and 70 species of protozoan parasites.[79] Some three-quarters of the links in food webs include a parasite, important in regulating host numbers. Perhaps 40 percent of described species are parasitic.[78]